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利用改进的GMA半薄连续切片技术对水稻品种IR36胚囊形成和发育进行了研究。结果表明：a）大孢子母细胞减数分裂Ⅰ形成的二分体和减数分裂Ⅱ形成的四分体的各个子细胞的大小都是不均等的；b）单核胚囊的3次有丝分裂的方向都较为固定；C）极核发育过程有明显的形态特征变化；d）胚囊形成各阶段淀粉粒出现与否有规律性。根据IR36胚囊形成的特点，初步将水稻胚囊形成过程分为8个时期，即孢原细胞形成期、大孢子母细胞形成期、大孢子母细胞减数分裂期、功能大孢子形成期、单核胚囊形成期、胚囊有丝分裂期、八核胚囊发育期和胚囊成熟期。
The formation and development of the rice embryo IR36 were studied by using the modified GMA semi-thin serial sectioning technique. The results showed that: a) meiosis of megasporocyte meiosis Ⅰ and the size of each of the four daughter cells formed by meiosis II were unequal; b) the third mitosis of monocytic embryo sac Are fixed in direction; C) There are obvious morphological changes in the development of polar nuclei; d) There are regular occurrence of starch grains in various stages of embryo sac formation. According to the formation of IR36 embryo sac, the process of embryo sac formation in rice was preliminarily divided into eight stages: the formation stage of sporoplasm, the formation of megaspore mother cells, the meiosis of megaspore mother cells, the formation of megaspores, Mononuclear embryo sac formation period, embryo sac mitosis, octopus embryo sac development and embryo sac maturation.