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The pollen wall protects pollen grains from abiotic and biotic stresses.During pollen wall development,tapetal cells play vital role by secreting proteins,signals and pollen wall material to ensure microspore development.But the regulatory mechanism underlying the secretory pathway of the tapetum is largely unknown.Here,we characterize the essential role of the Arabidopsis COPⅡ protein Secretory31B (SEC31B) in pollen wall development and secretive activity of tapetal cells.