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采用光学显微镜和透射电镜方法,研究管花肉苁蓉吸器侵入寄主根和幼苗分化建成过程的形态学。结果表明:1)管花肉苁蓉吸器接触柽柳根表皮层后,吸器顶端分化出侵入细胞。在侵入细胞引领下,吸器呈楔形结构依次侵入柽柳根皮层、中柱和木质部。在吸器侵入寄主根过程中,与寄主根皮层细胞接触的吸器细胞壁呈现增厚反应,与吸器顶端邻近的多层寄主皮层细胞被机械挤压而变形,乃至被破坏。2)在吸器侵入木质部后,与寄主根表皮连接的管花肉苁蓉芽端薄壁组织恢复分生能力,分化出芽端生长点;生长点分化鳞叶原基,鳞叶原基长大为鳞叶幼叶,在鳞叶幼叶的叶腋内分化出生长点,发育成肉苁蓉幼苗。结果显示,管花肉苁蓉吸器侵入寄主根是一个主动过程,并有机械力的参与。吸器侵入寄主根和芽端生长分别呈现单极生长方式。
Using optical microscopy and transmission electron microscopy methods, the morphological study of the invasion of C. tubulosa suction device into the root and seedling of the plant was established. The results showed that: 1) Cistanche tubulosum inhalation device contacted the tarsal root epidermis, the tip of the aspirate differentiated into cells. Invaded by cells in the lead, the suction device was wedge-shaped structure in turn invade tamarisk root cortex, the column and the xylem. In the process of invading the host into the host root, the cell wall of the aspirate that contacts with the host root cortex presents a thickening reaction, and the multi-layered host cortical cells adjacent to the apical end of the aspirator are deformed and even destroyed. 2) After the aspirator invades the xylem, the parenchyma of the tube of the flower buds of the flower buds connected with the host root epidermis restores the meristematic ability and differentiates into the bud-end growing point; the growing point differentiates the leaf primordium, Scale leaves leaves axillary differentiation of the growth point, developed into Cistanche seedlings. The results showed that the invasion of Cistanche tubulosum inhalation into the host root was an active process with the participation of mechanical force. Inhalation of the inhaler into the host root and shoot growth showed unipolar growth.