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在中国新疆和华南海相泥盆系中识别出交切、并列、叠覆遗迹组构（相关遗迹组构类）和欧氏、拓扑遗迹组构（本体遗迹组构类）．通过对火山碎屑风暴岩中复合交切遗迹组构的研究发现，共生组合在一起的遗迹化石，可划分出６个遗迹阶层，具有不同的先后、世代关系和指相意义．拓扑遗迹组构分析表明，庞大的雕画迹家族可归并为３个拓扑类，它们分别同胚于线、树、网．雕画迹从线至网的拓扑形态谱系结构，分别代表深水或缺氧环境中，造迹生物寻觅式、探索式和诱捕式的觅食行为．研究表明：遗迹组构分析对副层序、凝缩段、不同海平面变化速度形成的海进体系域和高水位体系域的识别和高频旋回的研究有重要作用
In the marine Devonian of Xinjiang and Huainan of China, the structures of intersections, juxtaposed and overlying relics (related relics) and the Euclidean and topological relics (ontology) are identified. Based on the study of the composition of the composite crosscutting vestiges in the volcaniclastic stormstones, it is found that the symbiotic assemblages of trace fossils can be divided into six relics, which have different precedents, generations and meanings. Topological ruins composition analysis shows that a large family of carvings and tracers can be merged into three topological categories, which are homeomorphic to lines, trees and nets respectively. The topological morphological structure of the sculptural trace from line to web represents the foraging, exploring and trapping foraging behavior of creatures in deep water or anoxic environment respectively. The results show that the structure analysis of ruins plays an important role in the identification and high-frequency cycle of the marine system tract and the highstand system tract formed by the secondary sequences, condensation segments and different sea-level velocities