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Reconstructing the origin of a polyploid species is particularly challenging when an ancestor has become extinct.Under such circumstances,the extinct donor of a genome found in the polyploid may be treated as a ghost species in that its prior existence is recognized through the presence of its genome in the polyploid.In this study,we aimed to determine the polyploid origin of Oxyria sinensis(2n = 40)for which onlyone congeneric species is known,that is diploid O.digyna(2n = 14).